David B. Lank


Department of Biological Sciences

Simon Fraser University

Burnaby, British Columbia

Canada V5A 1S6


(: 778-782‑3010                                  faxÊ: 778-782‑3496                            e‑:: dlank@sfu.ca                




    Born 13 August 1951, Rochester, New York. 

    U.S. and Canadian citizen


Web Site






     B.Sc.  Marlboro College, 1973.

     M.Sc. University of Minnesota, 1979, Dept. of Ecology and Behavioral Biology. 

     Ph.D.  Cornell University, 1983, Section of Neurobiology and Behavior.




     University Research Assoc. & Adj. Professor, Dept. Biol. Sci., Simon Fraser Univ., 1998–.

     Affiliate Faculty, Dept Biology and Wildlife, Univ. of Alaska Fairbanks, 2009-2010.

     Assistant Professor, limited term, Dept. of Biol. Sciences, Simon Fraser Univ., 1993–1998.

     Adjunct Professor, Department of Biology, Queen's University, 1991–1996.

     Research Associate, Department of Biology, Queen's University, 1984–1992.

     Postdoctoral Research Associate, Zoology Department, Ohio State Univ., 1984.

     Visiting Assistant Professor, Biology Department, Capital University, 1983.

     Instructor, Biology Department, University of North Dakota, 1982.

     Postdoctoral Research Fellow, Biology Dept, University of North Dakota, 1980–1981.


Research Interests


     Population consequences of individual decision making in animals, including relationships between animal behavior, ecology, population and conservation biology, and evolution.  My specific expertise is with the ecology of mating systems and migration in precocial birds.   


Teaching and Supervisory Experience


     Animal Behavior, Ornithology, Introductory Biology (team‑taught lectures, supervised

laboratories), Introductory Genetics (population genetics lectures), Population Biology

Seminars (Natural Selection, Evolution of Sex), Reproductive Strategies, Bird Migration Field Course.  Lecturer, NKE course in Alternative Reproductive Tactics. Erskine Fellow at Univ. of Canterbury, NZ.

     I have supervised both undergraduate and graduate thesis students.  I currently supervise 4 graduate students studying the behavioral ecology and conservation biology of shorebirds, and serve on additional graduate committees.  


Selected External Research Support

     NSERC Discovery grant:  Maintenance of Ecological Polymorphisms by Frequency-Dependent Selection (20142019).   

    SFU VP Research grant: Maintenance of polymorphic traits (2013).                                         

     NSF Polar Programs and Alaska Dept. Fish and Game: Sandpiper Population Biology in Changing Environments (with Brett Sandercock, Kansas State Univ): 2010–2014.

     DIREN (French Polynesia Env. Min.), USF&WS, Critical Environment Partnership Fund, Island Conservation: Conservation Biology of Tuamotu Sandpiper, 2008–2013.

     NSERC Discovery grants: "Sexual Selection and Genetic Polymorphism", 1995–2013.

     USF&WS: Neotropical Migratory Bird Conservation Act grant: Intrinsic Markers for

          Migratory Bird Conservation.  2008-2011.

     NSERC-SPG: Estuary Management and Sustainable Development. 2008–2011.

     NSERC-CRD, B.C. Forest Science Program, B.C. Min. of Forests, B.C. Forest Science Program, Coastal Forest Industry Consortium: “Habitat Selection by Marbled Murrelets”, 2002–2011.

     Seadoc Society: “Stable isotope analysis of Dunlin diet in the Skagit Estuary”, 2007.

     Canadian Wildlife Service: "Demography of Shorebirds", “Dunlin in BC”, 1994–2002.       

     Agriculture Canada, private NGOs: “Shorebird use of Agricultural Fields”, 1997–2000.

     NSERC: "Population Variation in Birds" (with Fred Cooke), 1992–1996.

     H.F. Guggenheim Foundation: "Behavioral Dimorphism in Ruff", 1989–91.

     NSF, Population Biology and Physiological Ecology Program:

            "Variability in Avian Mate Choice", 1985–87.

     National Geographic Society, Committee for Research and Exploration: 

           "Maintenance of Variation in Male Ruff", 1985, 1987, 1989.

     Fulbright Research Scholar in Finland, 1986.

     Netherlands–America Commission for Educational Exchange, 1986.



Chair, Simon Fraser University Northern Studies Committee: 2004.

Associate editor: Auk 2004–2008

.Reviewer for: Am. Nat., Ann. Rev. Ecol. Evol., Anim. Behav., Arctic, Ardea, Auk, B.C. Birds, Biol. Lett., Bird Behav., Behaviour, Behav. Ecol., Behav. Ecol. Sociobiol., Binat. Sci. Found., Biol. Conserv., Bird Behav., Can. J. Forest Res., Can. J. Zool., Condor, Conserv. Biol., Ecology, Evolution, Heredity, Horm. Behav., J. Anim. Ecol., J. Avian Biol. J. Field Ornithol, J. Wildl. Manage. NERC, NSF (Pop. Biol., Anim. Behav Psychobiol), NSERC (Evol. and Ecol), Northw. Sci., Ornis Fenn., Proc. Roy. Soc. Lond. B, Quart. Rev. Biol., Science, Wilson J. Ornithol.

Member: Canadian Marbled Murrelet Recovery Team, 2002

Organizer or co-organizer: Western Hemisphere Shorebird Group, Vancouver 2011; Society of Canadian Ornithologists' meeting, Vancouver 1999; 1–11th Western Sandpiper Workshops, 1994–2006: Vancouver and Victoria BC, Portland OR., Ensenada Mexico, Boulder CO.






English W.B., E. Kwon, B.K. Sandercock, D.B. Lank.  2017.  Effects of predator exclosures on reproductive success in Red-necked Phalaropes.  Wader Study (in press).


Brown, S., C. Gratto-Trevor, R. Porter, E.L. Weiser, D. Mizrahi, M. Boldenow, R. Bentzen, R. Clay, M.-A. Giroux, E.Kwon, D.B. Lank, N. Lecomte, J. Liebezeit, V. Loverti, J. Rausch, B.K. Sandercock, S. Schulte, P. Smith, A. Taylor, B. Winn, S. Yezerinac, and R.B. Lanctot.  2017.  Migratory connectivity of Semipalmated Sandpipers and implications for conservation.  Condor 119:207-224.  doi: 10.1650/CONDOR-16-55.1

Lank, D.B., C. Xu, B. Harrington, R.I.G. Morrison, C.L. Gratto-Trevor, P.W. Hicklin, B.K. Sandercock, P.A. Smith, E. Kwon, J. Rausch, L.D. Pirie Dominix, D.J. Hamilton, J. Paquet, S.E. Bliss, S. Neima, C. Friis, S.A. Flemming, A.M. Anderson and R.C. Ydenberg.  2017.  Multi-decade continent-wide increase and decrease in wing length, but not bill length, of a long-distance migratory shorebird.  Ecology and Evolution 7:3243-3256. doi: 10.1002/ece3.2898

Ydenberg R.C., Barrett J. Lank, D.B., Xu, C., Faber M.  2017. The redistribution of non-breeding dunlins in response to the post-DDT recovery of falcons.  Oecologia 183: 1101-1110. doi: 10.1007/s00442-017-3835-2

Bulla, M., M. Valcu, (13 authors), D.B. Lank, (59 authors) and B. Kempenaers. 2016.  Defying the 24h day: Unexpected diversity in socially synchronized rhythms of shorebirds.  Nature 540: 109113.  doi: 10.1038/nature20563

Tavera, E.A., D.B. Lank and P.M. González.  2016. Effects of migration distance on life history strategies of Western and Semipalmated Sandpipers in Perú.  Journal of Field Ornithology 87: 293–308.  doi:

Weiser, E.L., R.B. Lanctot, S.C. Brown, (24 authors), D.B. Lank, (20 authors) and B.K. Sandercock.  2016.  Effects of geolocators on hatching success, return rates, breeding movements, and change in body mass in 16 species of Arctic-breeding shorebirds. Movement Ecology 4:12.  doi: 10.1186/s40462-016-0077-6

Küpper, C., M. Stocks, J. E. Risse, N. dos Remedios, L.L. Farrell, S.B. McRae, T.C. Morgan, N. Karlionova, P. Pinchuk, Y.I. Verkuil, A.S. Kitaysky, J.C. Wingfield, T. Piersma, K. Zeng, J. Slate, M. Blaxter, D.B. Lank and T. Burke.  2016.  A supergene determines highly divergent male reproductive morphs in the ruff.  Nature Genetics 48: 79–83.  doi: 10.1038/ng.3443


Xu, C., J. Barrett, D.B. Lank and R.C. Ydenberg.  2015.  Large and irregular population fluctuations in migratory Pacific (Calidris alpina pacifica) and Atlantic (C. a. hudsonica) dunlins are driven by density-dependence and climatic factors.  Population Ecology 57: 551–567.  doi: 10.1007/s10144-015-0502-5

Taylor, C.M., D.B. Lank and B.K. Sandercock. 2015.  Using local dispersal data to reduce bias in apparent survival and mate fidelity.  Condor 117: 598–608.  doi: 10.1650/CONDOR-14-184.1

Farrell, L.L, C. Küpper, T. Burke and D.B.Lank.  2015.  Major breeding plumage color differences of male ruffs (Philomachus pugnax) are not associated with coding sequence variation in the melanocortin-1 receptor (MC1R) gene.  Journal of Heredity 106:211-215.  doi: 10.1093/jhered/esu079

English, W.B., D. Schamel, D.M. Tracy, D.F. Westneat and D.B. Lank.  2014.   Sex ratio varies with egg investment in the red-necked phalarope (Phalaropus lobatus).  Behavioral Ecology and Sociobiology 68:19391949.  doi: 10.1007/s00265-014-1800-1

Jamieson, S.E., R.C. Ydenberg and D.B. Lank.  2014.  Does predation danger on southward migration curtail parental investment by female western sandpipers?  Animal Migration 2:3443.  doi: 10.2478/ami-2014-0004

Verkuil, Y.I., J. C. Juillet, J. C.E.W. Hooijmeijer, D.B. Lank and T. Piersma.  2014.  Genetic variation in nuclear and mitochondrial markers supports a large sex difference in life-time reproductive skew in a lekking species.  Ecology and Evolution doi: 10.1002/ece3.1188

Thomson, R.L., V.-M. Pakanen, D. Tracy, L. Kvist, D.B. Lank, A. Rönkä and K. Koivula. 2014.  Providing parental care entails variable mating opportunity costs for male Temminck’s stints. Behavioral Ecology and Sociobiology 68:12611272.  doi: 10.1007/s00265-014-1737-4

Hope, D.D., D.B. Lank and R.C. Ydenberg.  2014.  Mortality-minimizing sandpipers vary stopover behaviour dependent on age and geographic proximity of migrating predators. Behavioral Ecology and Sociobiology 68:827-838.  doi: 10.1007/s00265-014-1695-x

Franks, S.E., D.B. Lank and W.H. Wilson.  2014.  Western Sandpiper.  The Birds of North America, Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology; http://bna.birds.cornell.edu/bna/species/090, doi: 10.2173/bna.90

Farrell, L.L., T. Burke, J. Slate, S.B. McRae and D.B. Lank.  2013. Mapping the female mimic morph locus on the microsatellite linkage map of the ruff.   BMC Genetics 14:109.  doi: 10.1186/1471-2156-14-109324

Publications (continued)

Franks S.E., G. Fernández, D.J. Hodkinson, T.K. Kyser and D.B. Lank.  2013.  The long and the short of it: no dietary specialisation between male and female western sandpipers despite strong bill size dimorphism.  PLoSOne doi:10.1371/journal.pone.0079835

Farrell, L.L., T. Burke, J. Slate and D.B. Lank.  2013. A first-generation microsatellite linkage map of the ruff.  Ecology and Evolution 3: 4631–4640. doi: 10.1002/ece3.830

Lank, D.B., L.L. Farrell, T. Burke, T. Piersma and S.B. McRae.  2013.  A dominant allele controls development into female mimic male and diminutive female ruffs.  Biology Letters 9: 20130653.  doi: 10.1098/rsbl.2013.0653

Hobson, K.A., G.L Slater, D.B. Lank, R.L.Milner and R. Gardiner.  2013.  Agricultural lands subsidize winter diet of Pacific dunlin Calidris alpina pacifica at two major estuaries.  Condor 115:515524.  doi: 10.1525/cond.2013.120118

Lozano, G.A. D.B. Lank and B. Addison.   2013. Immune and oxidative stress trade-offs in four classes of ruffs (Philomachus pugnax) with different reproductive strategies.  Canadian Journal of Zoology 91:212218.  doi: 10.1139/cjz-2012-0324

Nebel, S. D.M. Buehler, S. Kubli, D.B. Lank and C.C. Guglielmo.  2013.  Does innate immune function decline with age in captive ruffs Philomachus pugnax?  Animal Biology 63:233–240.  doi: 10.1163/15707563-00002409

Ekblom, R., L.L. Farrell, D.B. Lank and T. Burke.  2012.  Gene expression divergence and nucleotide differentiation between males of different colour morphs and mating strategies in the ruff.  Ecology and Evolution 2: 2485–2505. doi: 10.1002/ece3.370

Fernández G. and D.B. Lank.  2012.  Territorial behavior of Western Sandpipers on their nonbreeding grounds: effect of sex and foraging interference.  Journal of Field Ornithology 83:272281.   

Farrell, L.L., D.A. Dawson, G.J. Horsburgh, T. Burke and D.B. Lank.  2012.  Isolation, characterization and predicted genome locations of ruff (Philomachus pugnax, AVES) microsatellite loci.  Conservation Genetics Resources 4: 763–771.  doi: 10.1007/s12686-012-9639-0

Franks, S.E. D.R. Norris, T.K. Kyser, G. Fernández, B. Schwarz, R. Carmona, M.A. Colwell, J. Correa Sandoval, A. Dondua, H.R. Gates, B. Haase, D.J. Hodkinson, A. Jiménez, R.B. Lanctot, B. Ortego, B.K. Sandercock, F. Sanders J.Y. Takekawa, N. Warnock, R.C. Ydenberg and D.B. Lank.  2012.  Range-wide patterns of migratory connectivity in the western sandpiper Calidris mauri.  Journal of Avian Biology 155–167.

Gratto-Trevor, C.L., R.I.G. Morrison, D. Mizrahi, D.B. Lank, P. Hicklin and A.L. Spaans. 2012.  Migratory connectivity of Semipalmated sandpipers: Winter distribution and migration routes of breeding populations.  Waterbirds 35:8395.

van Rooyen, J., J.M. Malt and D.B. Lank.  2011.  Relating microclimate to epiphyte availability: edge effects on nesting habitat availability for the Marbled Murrelet.  Northwest Science 85:549–561.

Hope, D.D., D.B. Lank, B.D. Smith and R.C. Ydenberg.  2011. Migration of two calidrid sandpiper species on the predator landscape: how stopover time and hence migration speed vary with proximity to danger.  Journal of Avian Biology 42: 523530. 

        doi: 10.1111/j.1600-048X.2011.05347.x

Publications (continued)

Silvergieter, M.P. and D.B. Lank.  2011. Patch scale nest-site selection by Marbled murrelets (Brachyramphus marmoratus). Avian Conservation and Ecology – Écol. Conserve. oiseaux 6(2): 6. [online] URL: http://www.ace-eco.org/vol6/iss2/art6/

Silvergieter, M.P. and D.B. Lank.  2011.  Marbled murrelets select distinctive nest trees within old-growth forest patches.  Avian Conservation and Ecology – Écol. Conserve. oiseaux 6(2): 3. [online] URL: http://www.ace-eco.org/vol6/iss2/art3/

Jaatinen K., A. Lehikoinen and D.B. Lank.  2010.  Female-biased sex ratios and the proportion of cryptic male morphs of migrant juvenile ruffs (Philomachus pugnax) in Finland.  Ornis Fennica  87:125–134.

Fernández, G. and D.B. Lank.  2010.  Do sex and habitat differences in anti-predator behavior of Western Sandpipers Calidris mauri reflect cumulative or compensatory processes?  Journal of Ornithology 151:665–672.

Burger, A.E., R.A. Ronconi, M.P. Silvergieter, C.Conroy, V. Bahn, I.A. Manley, A. Cober and D.B. Lank.  2010.  Factors affecting the availability of thick epiphyte mats and other potential nest platforms for Marbled Murrelets in British Columbia.  Canadian Journal of  Forest Research 40:727–746.

Waterhouse F.L., A.E. Burger, P.K. Ott, A. Donaldson, D.B. Lank.  2010.   Does interpretation of Marbled Murrelet habitat change with different classification methods?  BC Journal of Ecosystems and Mangement 10:20–34.  www.forrex.org/publications/jem/ISS52/vol10_no3_art4.pdf

Ydenberg, R.C., D. Dekker, G. Kaiser, P.C.F. Shepherd, L.J. Evans Ogden, K. Rickards and D.B. Lank.  2010.  Winter body mass and over-ocean flocking as components of danger management by Pacific dunlins.  BMC Ecology 10:1. [online] URL:  www.biomedcentral.com/1472–6785/10/1.  doi: 10.1186/1472-6785-10-1

Malt J.M. and D.B. Lank. 2009.  Marbled Murrelet nest predation risk in managed forest landscapes: dynamic fragmentation effects at multiple scales.  Ecological Applications 19:1274–1287.

Burger, A.E., I.A. Manley, M. Silvergieter, D.B. Lank, K.M. Jordan, T.D. Bloxton and M.G. Raphael.  2009.  Re-use of nest sites by Marbled Murrelets (Brachyramphus marmoratus) in British Columbia. Northwest Naturalist 90:217–226.

Waterhouse, F.L., A.E. Burger, D.B. Lank, P.K. Ott, E.A. Krebs, and N. Parker.  2009.  Using the low-level aerial survey method to identify nesting habitat of Marbled Murrelets (Brachyramphus marmoratus). BC Journal of Ecosystems and Management 10:80–96.

Burger, A.E., F.L. Waterhouse, A. Donaldson, C. Whittaker and D.B. Lank.  2009.  New methods for assessing nesting habitat: air photo interpretation and low-level aerial surveys.  BC Journal of Ecosystems and Management 10:4–14.

Franks S.E., D.B. Lank, D.R. Norris, B.K. Sandercock, C.M. Taylor and T.K. Kyser.  2009.  Feather isotope analysis discriminates age-classes of Western, Least, and Semipalmated Sandpipers when plumage methods are unreliable. Journal of Field Ornithology 80:51–63.

 Zharikov Y., R.W. Elner, P.C.F. Shepherd and D.B. Lank.  2009.  Interplay between physical and predator landscapes affects transferability of shorebird distribution models.  Landscape Ecology 24: 129–144.  Doi: 10.1007/s10980-008-9291-y

Publications (continued)

Fernández, G. and D.B. Lank.  2008.  Foraging behaviour of non-breeding Western Sandpipers Calidris mauri as a function of sex, habitat and flocking.  Ibis 150: 518526.

Evans Ogden, L.J., S. Bittmann, D.B. Lank and F.C. Stevenson.  2008.  Factors influencing use of farmland habitat by shorebirds wintering in the Fraser River Delta, Canada. Agriculture, Ecosystems and Environment 124:252258.

 Taylor, C.M., D.B. Lank, A.C. Pomeroy and R.C. Ydenberg.  2008.  Relationship between stopover site choice of migrating sandpipers, their population status, and environmental stressors.  Israel Journal of Ecology and Evolution 53:245–261.

Waterhouse, F.L., A. Donaldson, D.B Lank, P.K. Ott and E.A Krebs.  2008.  Using air photos to interpret quality of Marbled Murrelet nesting habitat in south coastal British Columbia. BC Journal of Ecosystems and Management 9:17–37. www.forrex.org/publications/jem/ISS47/vol9_no1_art3.pdf

Evans Ogden, L.J., D.B. Lank and S. Bittmann.  2008.  A review of agricultural land use by shorebirds with special reference to habitat conservation in the Fraser River Delta, BC. Canadian Journal of Plant Science 88:71–83.

Fernández G. and D.B. Lank.  2008.  Effects of habitat loss on shorebirds during the non-breeding season: current knowledge and suggestions for action.  Ornitología Neotropical 19 (supp): 663–640.
Malt J.M. and D.B. Lank.  2007.  Temporal dynamics of edge effects on nest predation risk for the marbled murrelet.  Biological Conservation 140:160173.
Reneerkens, J., J.B. Almeida, D.B. Lank, J. Jukema, R.B. Lanctot, R.I.G. Morrison, W.I.C. Rijpstra, D. Schamel, H. Schekkererman, J.S. Sinninghe Damsté, P.S. Tomkovich, D.M. Tracy, I. Tulp and T. Piersma.  2007.  Parental role division predicts avian preen wax cycles.  Ibis 149:721–729.  doi: 10.1111/j.1474-919X.2007.00693.x

Ydenberg, R.C., R.W. Butler and D.B. Lank.  2007.  Effects of predator landscapes on the evolutionary ecology of routing, timing and molt by long-distance migrants.  Journal of Avian Biology 38:523–529.  doi: 10.1111/j.2007.0908-8857.04202.x

Fernández, G. and D.B. Lank.  2007.  Variation in the wing morphology of Western Sandpipers (Calidris mauri) in relation to sex, age-class, and annual cycle.  Auk 124:10371046.

Zharikov Y., D.B. Lank, F. Cooke.  2007.  Influence of landscape pattern on breeding distribution and success in a threatened Alcid, the marbled murrelet: model transferability and management implications.  Journal of Applied Ecology 44:748–759.

       doi: 10.1111/j.1365-2664.2007.01319.x

Norris, D.R., D.B. Lank, J. Pither, D. Chipley, R.C. Ydenberg and T.K. Kyser.  2007.  Trace element profiles as unique identifiers of western sandpiper (Calidris mauri) populations. Canadian Journal of Zoology 85:579583.

Lank, D.B. and Nebel S.  2006.  Cross-cutting research on a flyway scale – beyond monitoring.  In: Waterbirds Around the World.  G.C. Boere, C.A. Galbraith and D.A. Stroud, eds. pp. 107–112.  TSO Scotland Ltd., Edinburgh, UK.

Fernández, G. and D.B. Lank.  2006.  Sex, age, and body size distributions of Western Sandpipers during the nonbreeding season with respect to local habitat.  Condor 108:547557.

Publications (continued)

O’Hara, P.D., G. Fernández, B. Haase, H. de la Cueva and D.B. Lank.  2006.  Differential   migration of western sandpipers (Calidris mauri) with respect to body size and wing length. Condor 108:225232.

Zharikov, Y., D.B. Lank, F. Huettmann, R.W. Bradley, N. Parker, P. P.-W. Yen, L. McFarlane Tranquilla and F. Cooke.  2006.  Habitat selection and breeding success in a forest-nesting Alcid, the marbled murrelet, in two landscapes with different logging histories in south-western Canada.  Landscape Ecology 21:107120.  doi: 10.1007/s10980-005-1438-5

Evans Ogden, L.J., K.A. Hobson, D.B. Lank and S. Bittmann.  2005.  Stable isotope analysis reveals that agricultural habitat provides an important dietary component for nonbreeding Dunlin.  Avian Conservation and Ecology – Écol. Conserve. oiseaux 1: 3. [online] URL: http://www.ace-eco.org/vol1/iss1/art3/

McFarlane Tranquilla, L., N.R. Parker, R.W. Bradley, D.B. Lank, E.A. Krebs, L. Lougheed and C. Lougheed.  2005.  Breeding chronology of Marbled Murrelets varies between coastal and inshore sites in southern British Columbia.  Journal of Field Ornithology 76:357367.

Ydenberg, R.C., A. Niehaus and D.B. Lank.  2005.  Interannual differences in the relative timing of southward migration of  male and female western sandpipers (Calidris mauri). Naturwissenschaften 92:332335.

O’Hara, P.D., G. Fernández, F. Becerril, H. de la Cueva and D.B. Lank.  2005.  Life history varies with migratory distance in Western Sandpipers (Calidris mauri).  Journal of Avian Biology 36:191202.

Lozano, G.A. and D.B. Lank. 2004. Immunocompetence and testosterone-dependent condition traits in male ruffs (Philomachus pugnax).  Animal Biology. 54.315329.

Schamel, D., D.M. Tracy, D.B. Lank and D.F. Westneat.  2004.  Mate guarding, copulation strategies and paternity in the sex-role reversed, socially polyandrous Red-necked Phalarope Phalaropus lobatus. Behavioral Ecology and Sociobiology 57:110118.

Fernández, G., P.D. O’Hara and D.B. Lank.  2004.  Tropical and subtropical Western Sandpipers (Calidris mauri) differ in life history strategies.  Ornitología Neotropical 15 (suppl.) 385394.

Ydenberg, R.C., R.W. Butler, D.B. Lank, B.D. Smith and J. Ireland. 2004.  Western sandpipers have altered migration tactics as peregrine falcon populations have recovered.  Proceedings of the Royal Society of London, B 271:12631269.

Schamel, D., D.M. Tracy and D.B. Lank.  2004.  Male mate choice, male availability, and egg production as limitations on polyandry in the Red-necked Phalarope.  Animal Behaviour 67:847853.

Shepherd, P.C.F. and D.B. Lank.  2004.  Marine and agricultural habitat preferences of dunlin wintering in British Columbia.  Journal of Wildlife Management 68:6173.

Evans Ogden, L., K.A. Hobson and D.B. Lank   2004.  Blood isotopic (d13C and  d15N) turnover and diet-tissue fractionation factors in captive Dunlin.  Auk 121:170177.

Parker, N., E. Cam, D.B. Lank and F. Cooke.  2003.  Post-fledging survival of juvenile Marbled Murrelets as determined by radio-telemetry in Desolation Sound, British Columbia.  Marine Ornithology 31:207212.


Publications (continued)

McFarlane Tranquilla, L.A., P.-W. P. Yen, R.W. Bradley, B.A. Vanderkist, D.B. Lank, N.R. Parker, M. Drever, L.W. Lougheed, G. Kaiser and T.D. Williams.  2003.  Do two murrelets make a pair? Breeding status and behavior of Marbled Murrelet twosomes captured at-sea.  Wilson Bulletin 115:374381.

McFarlane Tranquilla, L.A., R. Bradley, N. Parker, D. Lank and F. Cooke. 2003.  Replacement laying in Marbled Murrelets.  Marine Ornithology 31:7581.

Lank, D.B., R.W. Butler, J. Ireland and R.C. Ydenberg.  2003.  Effects of predation danger on migration strategies of sandpipers.  Oikos 103:303–319.  doi: 10.1034/j.1600-0706.2003.12314.x

Lozano, G.A. and D.B. Lank.  2003.  Seasonal trade-offs in cell-mediated immunosenescence in ruffs (Philomachus pugnax).  Proceedings of the Royal Society of London, B 270:1203–1208.

Lank, D.B. and Ydenberg, R.C.  2003.  Death and danger at migratory stopovers: problems with “predation risk”.  Journal of Avian Biology 34:225228.

Fernández, G., H. de la Cueva, N. Warnock and D.B. Lank.  2003.  Apparent survival rates of Western Sandpiper (Calidris mauri) wintering in northwest Baja California, Mexico.  Auk 120:55–61.

McFarlane Tranquilla, L.A., R.W. Bradley, D.B. Lank, T.D. Williams, L.W. Lougheed and F. Cooke.  2003.  The reliability of brood patches in assessing reproductive status in the Marbled Murrelet: words of caution.  Waterbirds 26:108–118.

Lank, D.B.  2002.  Diverse processes maintain plumage polymorphisms in birds.  Journal of Avian Biology 33:327–330.

Nebel, S., D.B. Lank and 14 co-authors.  2002. Western Sandpipers (Calidris mauri) during the non-breeding season: spatial segregation on a hemispheric scale.  Auk 119:922–928.

Hazlitt S.L., R.C. Ydenberg and D.B. Lank.  2002.  Territory structure, parental provisioning, and chick growth in the Black Oystercatcher (Haematopus bachmani).  Ardea 90:219–227.

O’Hara, P.D., D.B. Lank and F.S. Delgado.  2002.  Is the timing of moult altered by migration? Evidence from a comparison of age and residency classes of Western Sandpipers (Calidris mauri) in Panamá.  Ardea 90:61–70.

Lank, D.B., C.M. Smith, O. Hanotte, A. Ohtonen, S. Bailey and T. Burke.  2002.  High frequency of polyandry in a lek mating system.  Behavioral Ecology 13:209–215.

Evans, M.R., D.B. Lank, W.S. Boyd, and F. Cooke.  2002.  The breeding ecology of Barrow's Goldeneye and Bufflehead nesting in nest boxes versus natural cavities.  Condor 104:610–619.

Ydenberg, R.C., R.W. Butler, D.B. Lank, C.G. Guglielmo M. Lemon and N. Wolf.  2002.  Trade-offs, condition dependence, and stopover site selection by migrating sandpipers.  Journal of Avian Biology 33:47–55.

Lank, D.B. and J. Dale.  2001.  Visual signals for individual identification: the silent "song" of Ruffs.  Auk 118:759–765.



Publications (continued)

Dale, J., D.B. Lank and H.K. Reeve.  2001.  Signaling individuality vs. quality: a model and case studies with ruffs, queleas and house finches.  American Naturalist 158:75–86.

Shepherd, P.C.F., D.B. Lank, B.D  Smith, N.D. Warnock, G.W. Kaiser and T.D. Williams.  2001.  Sex ratios of Dunlin wintering at two latitudes on the Pacific coast.

Condor 103:352–360.

Sandercock B.K., D.B. Lank, R.B. Lanctot, B. Kempenaers and F. Cooke.  2000.  Ecological correlates of mate fidelity in two arctic-breeding sandpipers.  Canadian Journal of  Zoology 78:1948–1958.

Lank, D.B., M. Coupe and K.E. Wynne-Edwards.  1999.  Testosterone-induced male traits in female ruffs (Philomachus pugnax): autosomal inheritance and gender differentiation.      Proceedings of the Royal Society of London, B 266:2323–2330.

Sandercock, B.K., D.B. Lank and F. Cooke.  1999.  Seasonal declines in the fecundity of arctic-breeding sandpipers: different tactics in two species with an invariant clutch size.                   Journal of Avian Biology 30:460–468.

Cooch, E.G., D.B. Lank, R.F. Rockwell and F. Cooke.  1999.  Body size and recruitment in snow geese.  Bird Study 46 (suppl.):S112–119.

Lougheed, L.W., A. Breault and D.B. Lank.  1999.  Estimating statistical power to evaluate ongoing waterfowl population monitoring.  Journal of Wildlife Management 63:1359–1369.

Lissimore, D., M.J.F. Lemon, D.B. Lank, R.W. Butler and R.C. Ydenberg.  1999.  Large and consistent body mass differences of migrant calidris sandpipers at adjacent stopover sites: phenomenon and possibilities.  Wader Study Group Bull. 88:55–58

Hugie, D.M. and D.B. Lank.  1997.  The resident's dilemma: a female‑choice model for the evolution of alternative male reproductive strategies in lekking male ruffs (Philomachus pugnax).  Behavioral Ecology 8:218–225.

Cooch, E.G., D.B. Lank, R.J. Robertson and F. Cooke.  1997.  Effects of parental age and environmental change on offspring sex-ratio in a precocial bird.  Journal of Animal Ecology 66:189–202.

Cooch, E.G., D.B. Lank and F. Cooke.  1996.  Intraseasonal variation in the development of sexual size dimorphism in a precocial bird.  Journal of Animal Ecology 65:439–450.

Lank, D.B., C.M. Smith, O. Hanotte, T.A. Burke and F. Cooke.  1995.  Genetic polymorphism for alternative mating behaviour in lekking male ruff,  Philomachus pugnax.                 Nature 378:59–62.

Robertson, G.J., E.G. Cooch, D.B. Lank, R.F. Rockwell and F. Cooke.  1994.  Female age and egg size in the lesser snow goose.  Journal of Avian Biology 25:149–155.

Downhower, J.F. and D.B. Lank.  1994.  Effect of previous experience on mate choice by female mottled sculpins.  Animal Behaviour 47:369–372.

Williams, T.D., D.B. Lank, F. Cooke and R.F. Rockwell.  1994.  Fitness consequences of egg‑size variation in lesser snow geese.  Oecologia 96:331–338.

Williams, T.D., D.B. Lank and F. Cooke.  1993.  Is intraclutch egg‑size variation adaptive in the lesser snow goose?  Oikos 67:250–256.


Publications (continued)

Lank, D.B. and C.M. Smith.  1992.  Females prefer larger leks: field experiments with ruffs (Philomachus pugnax).  Behavioral Ecology and Sociobiology 30:323–329.

Cooch, E.G., D.B. Lank, R.F. Rockwell and F. Cooke.  1992.  Is there a positive relationship      between body size and fecundity in Lesser Snow Geese?  Auk 109:667–673.

Lank, D.B., M.A. Bousfield, F. Cooke and R.F. Rockwell.  1991.  Why do snow geese adopt eggs?  Behavioral Ecology 2:181–187.

Cooke, F., R.F. Rockwell and D.B. Lank.  1991.  Recruitment in long‑lived birds: genetic considerations.  Proceedings of the 20th International Ornithological Congress:1666–1677.

Oring, L.W., J.M. Reed, M.A. Colwell, D.B. Lank and S.J. Maxson.  1991.  Factors regulating annual mating success and reproductive success in spotted sandpipers.   Behavioral Ecology and Sociobiology 28:433–442.

Cooch, E.G., D.B. Lank, R.F. Rockwell and F. Cooke.  1991.  Long-term decline in body size in a snow goose population: evidence of environmental degradation?  Journal of Animal Ecology 60:483–496.

Cooch, E.G., D.B. Lank, A. Dzubin, R.F. Rockwell and F. Cooke.  1991.  Body size variation in lesser snow geese: environmental plasticity in gosling growth rates.  Ecology 72:503–512.

Lank, D.B., R.F. Rockwell and F. Cooke.  1990.  Frequency‑dependent fitness consequences of intraspecific nest parasitism in snow geese.  Evolution 44:1436–1453. 

Lank, D.B., P. Mineau, R.F. Rockwell and F. Cooke.  1989.  Intraspecific nest parasitism and extra‑pair copulation in lesser snow geese Chen caerulescens.  Animal Behaviour 37:74–89.

Lank, D.B., E.G. Cooch, R.F. Rockwell and F. Cooke.  1989.  Environmental and demographic correlates of nest parasitism in lesser snow geese, Chen caerulescens caerulescens.              Journal of Animal Ecology 58:29–46.

Cooch, E.G., D.B. Lank, R.F. Rockwell and F. Cooke.  1989.  Long‑term decline in clutch size of a snow goose population: evidence for density‑dependence?  Journal of Animal Ecology 58:711–723.

Lank, D.B.  1989.  Why fly by night?  Inferences from tidally induced migratory departures of sandpipers.  Journal of Field Ornithology 60:154–161.

Lank, D.B. and C.M. Smith.  1987.  Conditional lekking in ruff (Philomachus pugnax).       Behavioral Ecology and Sociobiology 20:137–145.

Oring, L.W. and D.B. Lank.  1986.  Polyandry in spotted sandpipers: the impact of environment and experience.  In: Environmental Aspects of Social Evolution, D.I. Rubenstein and P. Wrangham, eds., pp. 21–41.  Princeton Univ. Press, Princeton.

Lank, D.B., L.W. Oring and S.J. Maxson.  1985.  Mate and nutrient limitation of egg‑laying in a polyandrous shorebird.  Ecology 66:1513–1524.

Oring, L.W. and D.B. Lank.  1984.  Breeding area fidelity, natal philopatry, and the social systems of sandpipers.  In: Shorebirds: Breeding Behavior and Populations, J. Burger and B.L. Olla, eds., pp. 125–147.  Plenum Press, NY.

Publications (continued)

Oring, L.W., D.B. Lank and S.J. Maxson.  1983.  Population studies of the polyandrous spotted sandpiper.  Auk 100:272–285.

Oring, L.W. and D.B. Lank.  1982.  Sexual selection, arrival times, philopatry, and site‑fidelity in the polyandrous Spotted Sandpiper.  Behavioral Ecology and Sociobiology 10:185–191.

Lank, D.B.  1979.  Dispersal and predation rates of wing‑tagged Semipalmated Sandpipers Calidris pusilla and an evaluation of the tag technique.  Wader Study Group Bulletin 27:41–46.




Cooke, F., R.F. Rockwell and D.B. Lank.  1995.  The Snow Geese of La Pérouse Bay: Natural Selection in the Wild.  Oxford University Press, Oxford.




Lank, D.B.  1983.  Migratory Behavior of Semipalmated Sandpipers at Inland and Coastal Staging Areas.  Ph.D. dissertation, Cornell University.

Lank, D.B.  1973.  Celestial Orientation by Migrating Sandpipers.  B.Sc. thesis, Marlboro College, Marlboro VT.



Overviews, letters, notes


Malt, J.M. and D.B. Lank  2007.  Observing wildlife in the canopies of old-growth trees using motion-sensitive cameras. Wildlife Afield 4:43-53.

Zharikov, Y., D.B. Lank, F. Huettmann and F. Cooke.  2007.  Interpreting habitat distribution models of an elusive species, the marbled murrelet: a response to Burger and Page.  Landscape Ecology 22:1283–1289.

Nebel, S. and D.B. Lank.  2003.  Cross-seasonal and cross-disciplinary interactions: Synergy from studying the year-round physiology, behaviour, and population biology of migratory shorebirds.  Wader Study Group Bulletin 100:118–121.

Butler, R.W., Ydenberg, R.C. and D.B. Lank. 2003.  Wader migration on the changing predator landscape.  Wader Study Group Buletin. 100:130–133.

Shepherd, P.C.F., L.J. Evans Ogden, and D.B. Lank.  2003.  Integrating marine and terrestrial habitats in shorebird conservation planning. Wader Study Group Bulletin 100:40–42.

Lank, D.B. and T. Piersma.  1989.  Ruffnet: the 1989 progress report. Wader Study Group Bulletin 54:34–35.

Lank, D.B. and T. Piersma.  1988.  Ruffnet: a ringing study of plumage and behavioral polymorphisms in Ruff.  Wader Study Group Bulletin 53:4–5.

LaDuke, J.C., D.B. Lank and T. Sirek.  1984.  Utilization of SAS as a revisionary tool and cataloguing program.  Assoc. Syst. Coll. Newslet.  12:13–14.




Marie-Hélène Burle, Caleb Ashling, Jean Durieux, Lucie Faulquier, Anne Gouni, Richard Lanctot, Sue McRae, Michel Poherui, Emmanuelle Portier, Pascal Roussel, Claude Serra and David Lank. 2009.   Etude du Chevalier des Tuamotu Prosobonia cancellatasur l’atoll de Tahanea: écologie, système social, reproduction et conséquences pour la conservation.  Ornithological Society of Polynesia – SOP-MANU.  www.manu.pf/PDF/Prosobonia_cancellata_2009.pdf.

Arcese P., A.E. Burger, C.L. Staudhammer,  J.P. Gibbs, E. Selak, G.D. Sutherland, J.D. Steventon, S.A. Fall, D. Bertram, I.A. Manley, S.E. Runyan, W.L. Harper, A. Harfenist, B.K. Schroeder, D.B. Lank, S.A. Cullen, J.A. Deal, D. Lindsay and G. Jones.  2005. Monitoring designs to detect population declines and identify their cause for the Marbled Murrelet.  Centre for Applied Conservation Research, University of British Columbia,

Fernández, G., N. Warnock, D.B. Lank and J.B. Buchanan.  2006. Conservation plan for the Western Sandpiper.  Manomet Center for Conservation Science, Manomet, Massachusetts.   Available online.

Waterhouse, F.L., A. Donaldson and D.B. Lank.  2004.  Using air photos to interpret Marbled Murrelet habitat in British Columbia: Application of a preliminary classification scheme.  Forest Research Technical Report TR-029, Coast Forest Region, Nanaimo, B.C., 37 pp.

Lank, D.B., N. Parker, E.A. Krebs and L. McFarlane Tranquilla.  2003.  Geographic distribution, habitat selection, and population dynamics with respect to nesting habitat characteristics of Marbled Murrelets Brachyramphus marmoratus.  Centre Wildlife Ecology, Simon Fraser University.  Available at: www.sfu.ca/biology/wildberg/species/mamu.html


Book and Meeting Reviews


Lank, D.B.  2014.  Reintroduction Biology.  ISBE Newslet. 26 (2): 8.

Ydenberg, R.C. and D.B. Lank. 2011.  Western Hemisphere Shorebird Group: Fourth meeting. Wader Study Group Bull. 1118:196-200. (meeting review).

Lank, D.B. 2011.  Shorebird Ecology, Conservation, and Management.  Condor 113: 703–704.

Lank,D.B.  2006.  Western Sandpiper Symposium.  Wader Study Group Bulletin 109:26. (meeting review).

Lank, D.B, K. Mathot, A. Pomeroy, R.C. Ydenberg.  2004  Behavioral Ecology surfaces in Finland: ISBE 2004.  ISBE Newslet. 16:913. (meeting review). 

Lank, D.B.  2004.  A Red Bird in a Brown Bag.  Quart. Rev. Biol. 79:87.      

Lank D.B.  1999.  Avian Growth and Development: Evolution within the Altricial-Precocial  Spectrum.  Quart Rev. Biol. 74:226–227.

Lank, D.B.  1998.  Partnership in Birds.  Int. Soc. Behav. Ecol. Newslet. 10:17–20.

Lank, D.B.  1997.  Foundations of Animal Behavior: Classic Papers with Commentaries.  Northwest Sci. 71:346–347.

Lank, D.B. and F.A. Pitelka.  1997.  “Leks” and “Arena Birds”.  Condor 99:558–559.

Lank, D.B.  1996.  The abundance and distribution of estuarine birds in the Strait of Georgia, British Columbia.  B.C. Birds  6:18–19.

Lank, D.B.  1994.  The Survival World of Birds.  Quart. Rev. Biol. 69:258.

Lank, D.B.  1991.  Homeward Bound.  Quart. Rev. Biol. 66:524.

Lank, D.B.  1986.  Bird Watching: State of the Art.  A review of: Current Ornithology, vol. 2.  Zoo Biol. 5:393–398.

Oring, L.W. and D.B. Lank.  1980.  Shorebirds in Marine Environments.  Auk 97:641–642.



Selected Published Abstracts


Lank, D.B., B. Crawford and C. Croton.  1998.  Behaviorally dimorphic male Ruffs have extremely long, but monomorphic, sperm.  Picoides 11(2):7.

Lank, D.B.  1993.  Migratory flight thresholds relative to estimated fat stores in Semipalmated Sandpipers (Calidris pusilla).  Abstract from American Ornithologists' Union meeting, 1993. Wader Study Group Bulletin 71:13–14.


Popular Scientific Articles


Küpper, C. and D.B. Lank.  2016.  Flipped genes don’t flip out.  Biosphere 13:27-30.


Selected Broadcast/Print Media


November 2015 (re: Nature Genetics study):

New Scientist: www.newscientist.com/article/dn28493-ruff-bird-orgies-have-four-sexes-thanks-to-a-supergene-flip/

As It Happens, CBC Radio, www.cbc.ca/radio/asithappens/as-it-happens-wednesday-edition-1.3324559/ruff-sex-scientists-identify-genetic-sequence-of-a-bird-with-four-genders-1.3324708;

CBC Radio 1 Kelowna; CKNW, Vancouver;  Radio Bangor (Wales)

Washington Post and syndicated papers, Canadian Press Story (Vancouver Sun, multiple Canadian newspapers)


Selected Presentations


Are Ruffs the most interesting birds in the world?  Plenary talk, Western Hemisphere Shorebird Group meeting III, Mazatlán, Mexico, March 2009, and COS-AOU-SCO meeting, San Diego CA Feb 2010.

Too Many Peregrines Cause Problems: Neglected Influence of Danger Management on Life History of their Prey.  Invited inaugural lecture for Katma award, Copper Ornithological Society, Arcata CA June 2005.  

Lek size and the equilibrium frequency of alternative male strategies in male ruffs. Int. Soc. Behav. Ecology, Jyväskylä, 2004  

Female choice in alternative male mating behaviour systems: overlooked or justifiably ignored?  Invited symposium, regional Anim. Behav. Soc., Columbia, MO, Mar. 2001.          

Lighter-weight migrant shorebirds indicate healthier ecosystems.  American Ornitologists' Union, St. John's, Aug. 2000. (with R.C. Ydenberg, R.W. Butler, K. O'Reilly, and J. Ireland).

First Incubation, hatching, captive rearing and release of a Marbled Murrelet (Brachyramphus marmoratus).  Society of Canadian Ornithologists, Montreal, 1999.

Do resident and satellite ruffs have dimorphic wintering strategies?  American Ornithologist' Union, 1999, Aug. 1999 (with C.M. Smith).

Behaviorally dimorphic male Ruffs have extremely long, but monomorphic, sperm.  Int. Soc. Behav. Ecology, July 1998 (with B. Crawford and C. Croton).

Behavioral mechanisms maintaining a genetic polymorphism in male mating strategy of ruffs. Invited talk, Evolution and Behavior of Birds, Brisbane, Australia, Sept. 1996

Old genes in new bodies: do shrinking Snow Geese reflect evolutionary change?  Invited symposium paper, Soc. for the Study of Evolution, Montreal, July 1995 (with E.G. Cooch and F. Cooke).

Sexual selection and behavioural polymorphisms in the Ruff.  Invited paper, Association for the Study of Animal Behaviour, London, 1993.

Migratory orientation and activity in sandpipers.  Wilson Ornithological Soc., 1979.

Stellar and magnetic orientation in sandpipers.  Symposium on Avian Orientation and

     Navigation, SUNY Albany, 1976.


Invited Seminars


Allegheny College

Bodega Bay Marine    


Canadian Wildlife       

      Service, B.C.

Carleton University

Cornell University

Eastern Michigan Univ.

Harvard University

McMaster University

Max Planck Institute Ohio State University Oregon State University

Queen's University

Simon Fraser University

Stanford University

SUNY Albany

Tulane University

Univ. of British Columbia

Univ. of Groningen, NL

Univ. of Leicester, U.K.

Univ. of Nevada, Reno

Univ. of North Carolina

     at Wilmington

Univ. of Oulu, Finland

University of Pennsylvania

Univ. of Richmond

Univ. Sheffield, UK

Univ. of Toronto  

Univ. of Uppsala, Sweden

Univ. of Washington

York University



Research Projects Supervised by David B. Lank


Thesis supervisor:


Schwarz, B.  2016.  Population structure in Western Sandpipers (Calidris mauri): variation in genes, morphology and vocalizations in a migratory shorebird.  Ph.D. thesis, Simon Fraser University.

Halpern, L.R.  2014.  Acoustic recorders reveal the impact of invasive rats on nocturnal burrow-nesting seabirds in Haida Gwaii.  M.REM project, Simon Fraser University. (with C. Berman).

English, W.E. 2014.  The evolutionary ecology of reproductive traits in the red-necked phalarope (Phalaropus lobatus).  M.Sc. thesis, Simon Fraser University.

Farrell, L.L.  2013. Molecular genetics of alternative reproductive morphs in ruffs.  Ph.D. thesis, Simon Fraser University.

Hentze N.T.  2012.  Characteristics of over-ocean flocking by Pacific dunlins (Calidris alpina pacifica) at Boundary Bay, British Columbia. M.Sc. thesis, Simon Fraser University. (with Ron Ydenberg).

Gardiner R.J.  2012.  Comparative stopover ecology of Least (Calidris minutilla) and Western (C. mauri) sandpipers during southward migration.  M.Sc. thesis, Simon Fraser University.

Franks S.E.  2012.  Population connectivity and the causes and consequences of differential migration in a long-distance migratory shorebird, the Western sandpiper.   Ph.D. thesis, Simon Fraser University.

Cortese L. 2011.  Picking Patches: What is the utility of habitat fragmentation in determining habitat use by local populations of the Marbled Murrelet, Brachyramphus marmoratus? : M.REM project, Simon Fraser University.

Morgan, T.  2010.  Hormonal regulation of alternative reproductive strategies.  M.Sc. thesis, University of Alaska, Fairbanks. (with A.S. Kitaysky). 

Hope, D.  2010.  The influence of the predator landscape on migratory decisions in two shorebird species. M.Sc. thesis, Simon Fraser University.  (with R. Ydenberg)

Jamieson, S.E.   2009.  Cross-seasonal factors affecting breeding investment by female Pacific Dunlins.  Ph.D. thesis, Simon Fraser University.  (with R. Ydenberg)

Silvergeiter, M.P. 2009.  Multi-scale analysis of nest site selection and fledging success by Marbled Murrelets (Brachyramphus marmoratus) in British Columbia.  M.Sc. thesis, Simon Fraser University.

Barrett, J.  2008.   The influence of oceanographic and terrestrial attributes on Marbled Murrelet Brachyramphus marmoratus) marine habitat selection during the breeding season. M.R.M.  research project, Simon Fraser University.

Malt, J.M.  2007.  The influence of habitat fragmentation on Marbled Murrelet (Brachyrampus marmoratus) habitat quality in southwestern British Columbia. M.Sc. thesis, Simon Fraser University.

Fernández, G.J.  2005.  Ecological and social factors affecting the local habitat distribution of Western Sandpipers wintering at Bahiá Santa María, northwest México. Ph.D. thesis, Simon Fraser University.

Nebel, S. 2003.  Ecological factors underlying the nonbreeding distribution of Western Sandpipers  Ph.D. thesis, Simon Fraser University, Burnaby, BC (with R. Ydenberg).

Evans, M.R.  2003.  Breeding habitat selection by Barrow’s Goldeneye and Bufflehead in the Cariboo-Chilcotin region of British Columbia: nest-sites, brood-rearing habitat, and competition.  Ph.D. thesis, Simon Fraser University. (with S. Boyd)

Evans Ogden, L.J.  2002.  Non-breeding shorebirds in a coastal agricultural landscape: winter habitat use and dietary sources.  Ph.D. thesis, Simon Fraser University.

O'Hara, P.  2002.  The role of feather wear in alternative life history strategies of a long-distance migratory shorebird, the Western Sandpiper (Calidris mauri). Ph.D. thesis, Simon Fraser University, Burnaby, BC

Shepherd, P.C.F.  2001.  Space use, habitat preferences, and time-activity budgets of non-breeding dunlin (Calidris alpina pacifica) in the Fraser River Delta, B.C.  Ph.D. thesis, Simon Fraser University.

Schamel, D.L.  2000.  Female and male reproductive strategies in the red-necked phalarope, a polyandrous shorebird.  Ph.D. thesis, Simon Fraser University.

Sandercock, B.K.  1997.  Factors affecting the breeding demography of Western Sandpipers (Calidris mauri) and Semipalmated Sandpipers (C. pusilla) at Nome, Alaska.  Ph.D. thesis, Simon Fraser University (with F. Cooke).

Pirie, A.J.  1996.  The energetic cost of lekking in male ruffs (Philomachus pugnax).  M.Sc. thesis, Queen’s University.

Marciano, M.T.  1993.  Criteria for female mate choice in the ruff (Philomachus pugnax).  B.Sc. Honours’ thesis, Queen’s University.

Sharma, N.B.  1992.  Inheritance of sex-limited plumage and behaviour in ruffs (Philomachus pugnax) – testosterone induced phenotype in females.  B.Sc. Honours’ thesis, Queen’s University.

Lougheed, L.W. 1991.  The roles of female choice and male-male competition in determining the mating success of male ruffs (Philomachus pugnax).  B.Sc. Honours’ thesis, Queen’s University.


Current graduate students:

Marie-Hélène Burle: Ph.D.  Conservation biology of endangered Tuamotu Sandpipers

Dave Hope: Ph.D.  Winter ecology of temperate sandpipers (with Ron Ydenberg)

Richard Johnson: Ph.D.  Winter ecology of tropical sandpipers (with Ron Ydenberg)

Eveling Tavera Fernandez: Ph.D.  Winter ecology of tropical sandpipers (with Ron Ydenberg)


Postdoctoral Fellows supervised

Natalie dos Remedios 2014: assortative mating with respect to relatedness

Cailin Xu 2013-2014: sandpiper population dynamics (with R. Ydenberg)

Who-Seung Lee 2010–2011:  Sex ratio allocation.

Caz Taylor 2005–2009: Individual-based modeling of sandpiper migration and demography

Yuri Zharikov 2005–2007: Non-breeding shorebird usage of estuaries.

                        2002–2003: Nesting habitat selection by Marbled Murrelets

Elsie Krebs 2003–2005: Marine and terrestrial home range use by Marbled Murrelets

Jim Dale 2001–2005: Signalling theory

George Lozano 1998–2001, 2009: Avian Immunoecology (with Ron Ydenberg)

Nils Warnock 1995–1996: Migration strategy of Western Sandpipers (with Lew Oring)


Technician/M.Sc. research associates supervised:

Skyla Witt (B.Sc) 2016: breeding captive sandpipers

Kathleen Cathcart (B.Sc) 2015: breeding captive sandpipers

Carlianne Dorman (B.Sc) 2014: breeding captive sandpipers

Michelle Martin (M.Sc.) 2013: breeding captive sandpipers

Teri Jones  (B.Sc) 2012-2013: breeding captive sandpipers, wildlife habitat modelling

Mel Hart: (Ph.D candidate) 2011: breeding captive sandpipers, DNA sexing

Ivy Whitehouse (M.Sc.) 2009: breeding biology of sandpipers

Ummat Somgee (B.Sc.) 2008: breeding biology of sandpipers

Nelli Ronka (B.Sc.) 2007.  Karyotypes of sandpipers

Nadine Parker (M.Sc) 2005.  Edge effects on Marbled Murrelet nests: an experimental test.

Jeanine Bond (M.Sc.) 2006.  Edge effects on Marbled Murrelet nests: an experimental test.

Laura McFarlane-Tranquilla (M.Sc.):  Breeding chronology and biology of Marbled Murrelets (2002-2005).

Melissa Mukai (B.Sc.):  Maintenance of polymorphism in breeding strategies 

Nadine Parker (M.Sc):  Post-breeding dispersal of Marbled Murrelets (2002-2004).


Graduate thesis committee member (selected)

Jeffery, Erica:  Intersexual dynamics of condition-dependent alternative male mating behaviours. Ph.D. (current)

Drake, A.E.G.  2013.  Seasonal interactions in migratory songbirds: assessing the strength of carry-over effects in the Yellow Warble (Dendroica petachina; aestive group).  Ph.D., Simon Fraser University.

Peterman, J. H.  2010: Maternal allocation decisions impacted by variable ecological conditions in a solitary bee: Behavioural ecology perspectives.  Ph. D. thesis, Simon Fraser University.

Whitehorne, I.  2009.  Differential overwinter survival of American Dippers in a partial migrant system. M.Sc., Simon Fraser University.

Yasué, M.  2006.  The breeding ecology and potential impacts of habitat change on the Malaysian plover Charadrius peronii in the Gulf of Thailand.  Ph.D. Univ. of Victoria.

Neihaus, A.  2003.  Factors influencing the southward migration timing of Western Sandpipers M.Sc. thesis, Simon Fraser University.   

Mitchell, J.  2003.  Queuing for reproduction in anemonefish. Ph.D. thesis, Simon Fraser University.

Moore, D.  2002. The provisioning tactics of parent Common terns (Sterna hirundo) in relation to brood energy requirement.  Ph.D. thesis, Simon Fraser University, Burnaby, BC.

Burns, J. 2001. The ecological consequences of fuel storage in migrating Calidris sandpipers. MSc thesis, Simon Fraser University, Burnaby, BC.

Wolf, N. 2001. Foraging ecology and site selection in Western Sandpipers during their fall migration through southwestern British Columbia. M.Sc. thesis, Simon Fraser University.

Willis, Pam  2001.  Mating system dynamics and hybridization of Dall’s (Phocoenoides dalli) and harbour porpoise (Phocoena phocoena).  M.Sc. thesis, Simon Fraser University.

Hazlitt, S.L.  1999.  Territory quality and parental behaviour of the black oystercatcher in the Strait of Georgia, British Columbia.  M.Sc. thesis, Simon Fraser University.