Western Sandpipers

Publications
on Shorebird Research

CWE logo
  Breeding Biology
Non-breeding Biology
Migration and Life Histories
Foraging Behaviour

Danger Management
Physiology
Fraser River Delta Projects
Other Shorebird Studies

Breeding Biology
Niehaus, A.C. and R.C. Ydenberg. 2006. Ecological factors associated with the breeding and migratory phenology of high-latitude breeding western sandpipers. Polar Biology 30: 11-17.
Niehaus, A.C., D.R. Ruthfauff and B.J. McCaffery. 2004. Response of predators to Western Sandpiper nest exclosures. Waterbirds 27: 79-82.
Nebel, S. and B.J. McCaffery. 2003. Vocalisation activity of breeding shorebirds: documentation of its seasonal decline and applications for bird surveys. Can. J. Zool. 81: 1702-1708.
Niehaus, A.C. 2003. Ecology of migratory timing by southbound male and female western sandpipers (Calidris mauri). MSc, Simon Fraser University, Burnaby, BC. 127 pp.
Sandercock, B.K., D.B. Lank, R.B. Lanctot, B. Kempenaers and F. Cooke. 2000. Ecological correlates of mate fidelity in two arctic-breeding sandpipers. Can. J. Zool. 78: 1948-1958.
Sandercock, B.K., D.B. Lank and F. Cooke. 1999. Seasonal declines in fecundity of arctic-breeding sandpipers: different tactics in two species with an invariant clutch size. J. Avian Biol. 30: 460-468.
Sandercock, B. 1998. Assortative mating and sexual size dimorphism in Western and Semipalmated Sandpipers. Auk 115: 786-790.
Sandercock, B. 1998. Chronology of nesting events in Western and Semipalmated sandpipers near the Arctic Circle. J. Field Ornithol. 69: 235-243.
Lanctot, R.B., B.K. Sandercock and B. Kempenaers. 1997. Breeding displays in an arctic wader: who does them, and how display intensity relates to mate acquisition and nesting success. Advances in Ethology 32:212.
Sandercock, B. 1997. Incubation capacity and clutch size determination in two calidrine sandpipers: a test of the four-egg threshold. Oecologia 110: 50-59.
Sandercock, B.K. 1997. Factors affecting the breeding demography of Western Sandpipers (Calidris mauri) and Semipalmated Sandpipers (C. pusilla) at Nome, Alaska. PhD, Simon Fraser University, Burnaby, BC. 134 pp.
Sandercock, B.K. and C.L. Gratto-Trevor. 1997. Local survival in Semipalmated Sandpipers Calidris pusilla breeding at La Perouse Bay, Canada. Ibis 139: 305-312.
Sandercock, B. 1996. Removal of eggshells at hatch by Western and Semipalmated sandpipers. Condor 98: 431-433.

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Non-breeding Biology, including Panama and Mexico
Fernández, G.J. and D.B. Lank. Submitted. Sex differences of non-breeding western sandpipers (Calidris mauri) in foraging behavior and susceptibility to interference. Ibis.
Fernandez, G.J. and D.B. Lank. 2006. Sex, age, and body size distributions of Western Sandpipers during the nonbreeding season with respect to local habitat. Condor 108: 547-557.
Fernandez Aceves, G.J. 2005. Ecological and social factors affecting the local habitat distribution of Western Sandpipers wintering at Bahia Santa Maria, northwest Mexico. PhD, Simon Fraser University, Burnaby, BC. 117 pp.
Robinson, R.A., N.A. Clark, R. Lanctot, S. Nebel, B. Harrington, J.A. Clark, J.A. Gill, H. Meltofte, D.I. Rogers, K.G. Rogers, B.J. Ens, C.M. Reynolds, R.M. Ward, T. Piersma and P.W. Atkinson. 2005. Long term demographic monitoring of wader populations in non-breeding areas. Wader Study Group Bull. 106:17-29.
Fernandez, G., P.D. O'Hara and D.B. Lank. 2004. Tropical and subtropical Western Sandpipers (Calidris mauri) differ in life history strategies. Ornit. Neotrop. 15 (Suppl.): 385-394.
Fernandez, G., H. de la Cueva, N. Warnock and D.B. Lank. 2003. Apparent survival rates of Western Sandpipers (Calidris mauri) wintering in northwest Baja California, Mexico. Auk 120: 55-61.
Danemann, G.D., R. Carmona and G. Fernandez. 2002. Migratory shorebirds in the Geurrero Negro Saltworks, Baja California Sur, Mexico. Wader Study Group Bull. 97:36-41.
Nebel, S., D.B. Lank, P.D. O'Hara, G. Fernandez, B. Haase, F. Delgado, F.A. Estela, L.J. Evans Ogden, B. Harrington, B.E. Kus, J.E. Lyons, F. Mercier, B. Ortego, J.Y. Takekawa, N. Warnock and S.E. Warnock. 2002. Western Sandpipers (Calidris mauri) during the non-breeding season: spatial segregation on a hemispheric scale. Auk 119: 922-928.
Fernandez, G., H. de la Cueva and N. Warnock. 2001. Phenology and length of stay of transient and wintering Western Sandpipers at Estero Punta Banda, Mexico. J. Field Ornithol. 72: 509-520.
Buenrostro, M.A., N. Warnock and H. de la Cueva. 1999. Wintering Western Sandpipers (Calidris mauri) at Estero de Punta Banda, Baja California, Mexico. Wader Study Group Bulletin 88:59-63.
Becerril, F. 1998. Interaccion ecologica: conducta de forrajeo y territorialidad entre Calidris mauri y Calidirs minutilla (Charadriiformes: Scolopacidae) en el estero Punta Banda (B.C., Mexico), en la temporada invernal de 1996-1997. Thesis, CICESE, pp.
Butler, R.W., R.I.G. Morrison, R.S. Delgado, R.K. Ross and G.E.J. Smith. 1998. Distribution of shorebirds, coastal seabirds and wading birds in relation to oceanic upwelling along the Pacific coast of Panama. In: Atlas of Nearctic shorebirds and other waterbirds on the coast of Panama, Morrison, R.I.G., R.W. Butler, F.S. Delgadoand R.K. Ross, eds., pp. 90-103. Ottawa: Canadian Wildlife Service Special Publications.
Fernandez, G., R. Carmona and H. de la Cueva. 1998. Abundance and distribution of western sandpipers in Ensenada de La Paz, Baja California Sur. Southwest. Nat. 43: 57-61.
Morrison, R.I.G., F.S. Delgado, R.K. Ross and R.W. Butler. 1998. The distribution and abundance of Nearctic shorebirds on the coast of Panama. In: Atlas of Nearctic shorebirds and other waterbirds on the coast of Panama, Morrison, R.I.G., R.W. Butler, F.S. Delgadoand R.K. Ross, eds., pp. 41-68. Ottawa: Canadian Wildlife Service Special Publications.
Butler, R.W., R.I.G. Morrison, K.R. Ross, F.S. Delgado and G.E.J. Smith. 1997. Habitat associations of coastal birds in Panama. Colonial Waterbirds 20: 518-524.
Warnock, N., G.W. Page and B.K. Sandercock. 1997. Local survival of Dunlin wintering in California. Condor 99: 906-915.
Buenorostro, A. 1996. Estructura de la poblacion y uso de habitat de Calidris mauri en la temporada invernal de 1994-1995 en estero Punta Banada, Ensenada, Baja California, Mexico. Thesis, CICESE, Ensenada, Mexico, pp.
Fernandez, G. 1996. Tasas de retorno y residencia de Calidris mauri (Charadriiformes: Scolopacidae) en el estero Punta Banda (B.C., Mexico) en la temporada invernal de 1995-1996. MSc, Centro de Investigación Científica de Educación Superior de Ensenada (CICESE), Ensenada. pp.
Sandoval, Y. 1996. Relacion de la masa con la condicion corporal de Calidris mauri (Aves: Scolopacidae) en estero Punta Banda, durante el invierno 1994-1995. Thesis, CICESE, Ensenada. pp.

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Migration and Life Histories
Fernández, G. and D.B. Lank. 2007. Variation in the wing morphology of Western Sandpipers (Calidris mauri) in relation to sex, age class, and annual cycle. Auk 124: 1037-1046.
Lank, D.B. and S. Nebel. 2007. Cross-cutting research on a flyway scale - beyond monitoring. In: Waterbirds Around the World, Boere, G.C., C.A. Galbraithand D.A. Stroud, eds., pp. 107-112. Edinburgh: TSO Scotland Ltd.
Mathot, K., B.D. Smith and R.W. Elner. 2007. Latitudinal clines in food distribution correlate with differential migration in the western sandpiper. Ecology 88: 781-791.
Norris, D.R., D.B. Lank, J. Pither, D. Chipley, R.C. Ydenberg and T.K. Kyser. 2007. Trace element profiles as unique identifiers of western sandpiper (Calidris mauri) populations. Can. J. Zool. 85: 579-583.
Taylor, C.M. and D.R. Norris. 2007. Predicting conditions for migration: effects of density dependence and habitat quality. Biol. Let. 3: 280-283.
Nebel, S. 2006. Latitudinal clines in sex ratio, bill, and wing length in Least sandpipers. J. Field Ornithol. 77: 39-45.
Norris, D.R. and C.M. Taylor. 2006. Predicting the consequences of carry-over effects on migratory populations. Biol. Let. 2: 148-151.
O'Hara, P.D., G. Fernandez, B. Haase, H. de la Cueva and D.B. Lank. 2006. Differential migration of Western Sandpipers (Calidris mauri) with respect to body size and wing length. Condor 108: 225-232.
Nebel, S. 2005. Latitudinal clines in bill length and sex ratio in a migratory shorebird: a case of resource partitioning? Act. Oecolog. 28: 33-38.
O'Hara, P.D., G. Fernandez, F. Becerril, H. de la Cueva and D.B. Lank. 2005. Life history varies with migratory distance in Western Sandpipers (Calidris mauri). J. Avian Biol. 36: 191-202.
Ydenberg, R.C., A.C. Niehaus and D.B. Lank. 2005. Interannual differences in the relative timing of southward migration of male and female western sandpipers (Calidris mauri). Naturwissensch. 92: 332-335.
Nebel, S., A. Cloutier and G.J. Thompson. 2004. Molecular sexing of prey remains permits a test of sex-biased predation in a wintering population of western sandpipers. Proc. Roy. Soc. Lond. (Suppl.) 271: S321-S323.
Burns, J.G. 2003. Relationship of Calidris sandpiper wing shape with relative fuel load and total migration distance. Auk 120: 827-835.
Nebel, S. 2003. Ecological factors underlying the non-breeding distribution of western sandpipers. PhD, Simon Fraser University, Burnaby. 96 pp.
Nebel, S. and D.B. Lank. 2003. Cross-seasonal and cross-disciplinary interactions: synergy from studying the year-round physiology, behaviour, and population biology of migratory shorebirds. Wader Study Group Bull. 100:118-121.
Shepherd, P.C.F., L.J. Evans Ogden and D.B. Lank. 2003. Integrating marine and terrestrial habitats in shorebird conservation planning. Wader Study Group Bulletin 100:40-42.
O'Hara, P. 2002. The role of feather wear in alternative life history strategies of a long-distance migratory shorebird, the Western Sandpiper (Calidris mauri). PhD, Simon Fraser University, Burnaby, BC. 173 pp.
O'Hara, P.D., D.B. Lank and F.S. Delgado. 2002. Migration and wing feather moult trade-off in Western Sandpipers (Calidris mauri) wintering in Panama. Ardea 90: 61-70.
Butler, R.W., N.C. Davidson and I.G. Morrison. 2001. Global scale shorebird distribution in relation to productivity of near-shore ocean waters. Waterbirds 24:224-232.
Morbey, Y.E. and R.C. Ydenberg. 2001. Protandrous arrival timing to breeding areas: a review. Ecol. Letters 4: 663-673.
Shepherd, P.C.F., D.B. Lank, B.D. Smith, N. Warnock, G.W. Kaiser and T.D. Williams. 2001. Sex ratios of Dunlin wintering at two latitudes on the Pacific coast. Condor 103: 352-360.
Wolfe, N. 2001. Foraging ecology and site selection in Western Sandpipers during their fall migration through southwestern British Columbia. MSc, Simon Fraser University, Burnaby, BC. 118 pp.
Butler, R.W., T.D. Williams, N. Warnock and M.A. Bishop. 1999. Reply to Hedenstrom and Weber's "Gone with the wind? A comment on Butler et al." Auk 116: 563-564.
Clark, C.W. and R.W. Butler. 1999. Fitness components of avian migration: a dynamic model of  Western Sandpiper migration. Evol. Ecol. Rev. 1: 443-457.
Butler, R.W., T.D. Williams, N. Warnock and M.-A. Bishop. 1997. Wind assistance: a requirement for migration of small shorebirds? Auk 114: 456-466.
Butler, R.W., F.S. Delgado, H. de la Cueva, V. Pulido and B.K. Sandercock. 1996. Migration routes of the Western sandpiper. Wilson Bull. 108: 662-672.
Iverson, G.C., S. Warnock, R. Butler, M.A. Bishop and N. Warnock. 1996. Spring migration of Western Sandpipers along the Pacific coast of North America: a telemetry study. Condor 98: 10-21.
Lank, D.B. 1996. The abundance and distribution of estuarine birds in the Strait of Georgia, British Columbia. B. C. Birds 6:18-19.

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Foraging Behaviour
Stein, R.W., G.J. Fernández, H. de la Cueva and R.W. Elner. Submitted. Disproportionate bill length dimorphism and niche differentiation in wintering western sandpipers (Calidris mauri). Can. J. Zool.
Kuwae, T., P. Beninger, P. Decottignies, K.J. Mathot, D.R. Lund and R.W. Elner. 2008. Biofilm grazing in a higher vertebrate: the western sandpiper, Calidris mauri. Ecology 89:599-606. (Video clip of sandpiper feeding)
Mathot, K.J. 2005. Sex-related differences in feeding behaviour and implications for differential migration in Western Sandpipers (Calidris mauri). MSc, Simon Fraser University, Burnaby, BC. 102 pp.
Nebel, S., D.L. Jackson and R.W. Elner. 2005. Functional association of bill morphology and foraging behaviour in Calidrid sandpipers. Anim. Biol. 55: 235-243.
Nebel, S. and G.J. Thompson. 2005. Foraging behaviour of Western Sandpipers changes with sediment temperature:  implications for their hemispheric distribution. Ecol. Res. 20: 503-507.
Pomeroy, A.C. and R.W. Butler. 2005. Colour infrared photography is not a good predictor of macro invertebrate abundance on mudflats used by shorebirds. Waterbirds 28: 1-7.
Mathot, K.J. and R.W. Elner. 2004. Evidence for sexual partitioning of foraging mode in western sandpipers (Calidris mauri) during migration. Can. J. Zool. 82: 1035-1042.
Elner, R.W. and D. Seaman. 2003. Calidrid conservation: unrequired needs. Wader Study Group Bull. 100:30-34.
Sutherland, T., P.C.F. Shepherd and R.W. Elner. 2000. Predation on meiofaunal and macrofaunal invertebrates by Western Sandpipers (Calidris mauri):  evidence for selection and dual feeding modes. Marine Biol. 137: 983-993.
Shepherd, P. and Boates. 1999. Effects of a commercial baitworm harvest on Semipalmated Sandpipers (Calidris pusilla) and their prey in the Bay of Fundy Hemisphere Shorebird Reserve. Conservation Biology 13: 347-356.
Backwell, P.R.Y., P.D. O'Hara and J.H. Christy. 1998. Prey availability and selective foraging in shorebirds. Anim. Behav. 55: 1659-1667.

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Danger Management
Pomeroy, A.C., D.A. Acevedo Seaman, R.W. Butler, R.W. Elner, T.D. Williams and R.C. Ydenberg. In revision. Feeding-danger tradeoffs underlie stopover site selection by migrants. Anim. Behav.
Ydenberg, R.C., R.W. Butler and D.B. Lank. 2007. Effects of predator landscapes on the evolutionary ecology of routing, timing and molt by long-distance migrants. J. Avian Biol. 38: 523-529.
Pomeroy, A.C. 2006. Feeding and predation danger tradeoffs in stopover site usage by Western Sandpipers (Calidris mauri). PhD, Simon Fraser University, Burnaby. pp.
Pomeroy, A.C. 2006. Trade-offs between food abundance and predation danger in spatial usage of a stopover site by western sandpipers, Calidris mauri. Oikos 112: 629-637.
Pomeroy, A.C., R.W. Butler and R.C. Ydenberg. 2006. Experimental evidence that migrants adjust usage at a stopover site to trade off food and danger. Behav. Ecol. 17: 1041-1045.
Nebel, S. and R.C. Ydenberg. 2005. Differential predator escape performance contributes to a latitudinal sex ratio cline in a migratory shorebird. Beh. Ecol. Sociobiol. 59: 44-50.
Dekker, D. and R.C. Ydenberg. 2004. Raptor predation on wintering dunlins in relation to the tidal cycle. Condor 106: 415-419.
Ydenberg, R.C., R.W. Butler, D.B. Lank, B.D. Smith and J. Ireland. 2004. Western sandpipers have altered migration tactics as peregrine falcon populations have recovered. Proc. Roy. Soc. Lond. B. 271: 1263-1269.
Butler, R.W., R.C. Ydenberg and D.B. Lank. 2003. Wader migration on the changing predator landscape. Wader Study Group Bull. 100:130-133.
Lank, D.B., R.W. Butler, J. Ireland and R.C. Ydenberg. 2003. Effects of predation danger on migratory strategies of sandpipers. Oikos 103: 303-319.
Lank, D.B. and R.C. Ydenberg. 2003. Death and danger at migratory stopovers: problems with "predation risk". J. Avian Biol. 34: 225-228.
Burns, J.G. and R.C. Ydenberg. 2002. The effects of wing loading and gender on the escape flights of Least Sandpipers (Calidris minutilla) and Western Sandpipers (Calidris mauri). Beh. Ecol. Sociobiol. 52: 128-136.
Ydenberg, R.C., R.W. Butler, D.B. Lank, C.G. Guglielmo, M. Lemon and N. Wolf. 2002. Trade-offs, condition dependence, and stopover site selection by migrating sandpipers. J. Avian Biol. 33:47-55.
Burns, J. 2001. The ecological consequences of fuel storage in migrating Calidris sandpipers. MSc, Simon Fraser University, Burnaby, BC. 88 pp.
Burns, J.G. and S.L. Wardrop. 2001. The veloci-raptor:  a bicycle-powered model raptor for realistic predator encounter experiments. J. Field Ornithol. 72: 399-403.
Lissimore, D., M.J.F. Lemon, D.B. Lank, R.W. Butler and R.C. Ydenberg. 1999. Large and consistent body mass differences of migrant Calidris sandpipers at adjacent stopover sites: phenomenon and possibile explanations. Wader Study Group Bull. 88:55-58.

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Physiology
Reneerkens, J., J.B. Almeida, D.B. Lank, J. Jukema, R.B. Lanctot, R.I.G. Morrison, W.I.C. Rijpstra, D. Schamel, H. Schekkererman, J.S. Sinninghe Damsté, P.S. Tomkovich, D.M. Tracy, I. Tulp and T. Piersma. Submitted. Parental role division predicts avian preen wax cycles: a comparative study of sandpipers. Ibis.
Williams, T.D., N. Warnock, J.Y. Takekawa and M.A. Bishop. 2007. Flyway scale variation in plasma triglyceride levels as an index of refueling rate in spring migrating Western Sandpipers (Calidris mauri). Auk 124: 886-897.
Acevedo Seaman, D.A., C.G. Guglielmo, R.W. Elner and T.D. Williams. 2006. Landscape-scale physiology: Site differences in refueling rates indicated by plasma metabolite analysis in free-living migratory sandpipers. Auk 123: 563-574.
Stein, R.W. and T.D. Williams. 2006. Causes and consequences of post-growth age-dependent differences in small intestine size in a migratory sandpiper (Calidris mauri, Western Sandpiper). Funct. Ecol. 20: 142-150.
Seaman, D.A., C.G. Guglielmo and T.D. Williams. 2005. Effects of physiological state, mass change, and diet on plasma metabolite profiles in the western sandpiper (Calidris mauri). J. Exp. Biol. 208: 761-769.
Stein, R.W., A.R. Place, T. Lacourse, C.G. Guglielmo and T.D. Williams. 2005. Digestive organ sizes and enzyme activities of refueling western sandpipers (Calidris mauri): contrasting effects of season and age. Physiological and Biochemical Zoology 78: 434-446.
Egeler, O., D. Seaman and T.D. Williams. 2003. Influence of diet on fatty-acid composition of depot fat in Western Sandpipers (Calidris mauri). Auk 120: 337-345.
Guglielmo, C.G. and T.D. Williams. 2003. Phenotypic flexibility of body composition in relation to migratory state, age and sex in the Western Sandpiper (Calidris mauri). Physiol. Biochem. Zool. 76: 84-98.
Seaman, D.A. 2003. Landscape physiology: plasma metabolites, fattening rates and habitat quality in migratory western sandpipers. MSc, Simon Fraser University, Burnaby, BC. 114 pp.
Stein, R.W. and T.D. Williams. 2003. Tissue damage precludes the use of the everted sleeve technique to measure nutrient uptake in a small migratory shorebird, the Western Sandpiper (Calidris mauri). Physiol. Biochem. Zool. 76: 762-770.
Guglielmo, C.G., N.H. Haunerland, P.W. Hochachka and T.D. Williams. 2002. Seasonal dynamics of flight muscle fatty acid binding protein and catabolic enzymes in a migratory shorebird. Amer. J. Physiol. 282: R1405-1413.
Guglielmo, C.G., P.D. O'Hara and T.D. Williams. 2002. Extrinsic and intrinsic sources of variation in plasma lipid metabolites in free-living Western Sandpipers (Calidris mauri). Auk 119: 437-445.
Guglielmo, C.G., T.D. Williams, G. Zwingelstein, G. Brichon and J.-M. Weber. 2002. Plasma and muscle phospholipids are involved in the metabolic response to long-distance migration in a shorebird. Journal of Comparative Physiology B 172:409-417.
McFarland, C.N., L.I. Bendell-Young, C. Guglielmo and T.D. Williams. 2002. Kidney, liver and bone cadmium content in the Western Sandpiper in relation to migration. J. Environ. Monit. 4: 791-795.
Stein, R.W. 2002. Busting a gut: Age-related variation and seasonal modulation of digestive tract structure and function in the Western Sandpiper. MSc, Simon Fraser University, Burnaby, BC. 114 pp.
Guglielmo, C.G. and J.G. Burns. 2001. Avian forensics: predicting body fat and body mass from wing remains. J. Avian Biol. 32: 198-203.
Guglielmo, C.G., T. Piersma and T.D. Williams. 2001. A sport physiological perspective on bird migration: evidence for flight-induced muscle damage. J. Exp. Biol. 204: 2683-2690.
Egeler, O. and T.D. Williams. 2000. Seasonal, age and sex-related variation in fatty-acid composition of depot fat in relation to migration in the Western Sandpiper. Auk 117: 110-119.
Egeler, O., T.D. Williams and C.G. Guglielmo. 2000. Modulation of lipogenic enzymes, fatty acid synthase and D9-desaturase, in relation to migration in the Western Sandpiper (Calidris mauri). Journal of Comparative Physiology B 170: 169-174.
Guglielmo, C.G. 1999. Physiological and biochemical modulation for long-distance migration:  the functional significance of intra-specific variation in the Western Sandpiper. PhD, Simon Fraser University, Burnaby, BC. 226 pp.
Williams, T.D., C.G. Guglielmo, O. Egeler and C.J. Martyniuk. 1999. Plasma lipid metabolites provide information on mass change over several days in captive Western Sandpipers. Auk 116: 994-1000.
Egeler, O. 1998. Aspects of fatty acid synthesis and composition in relation to migration in the Western Sandpiper (Caldris mauri). MSc, Simon Fraser University, Burnaby, BC. 99 pp.
Guglielmo, C.G., N.H. Haunerland and T.D. Williams. 1998. Fatty acid binding protein, a major protein in the flight muscle of migrating Western Sandpipers. Comp. Biochem. Physiol. 119: 549-555.

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Fraser River Delta Projects
Evans Ogden, L.J., S. Bittmann, D.B. Lank and F.C. Stevenson. Submitted. Factors influencing use of farmland habitat by shorebirds wintering in the Fraser River Delta, Canada. Agric., Ecosys. Environ.
Evans Ogden, L.J., D.B. Lank and S. Bittmann. In press. Use of agricultural land by shorebirds with special reference to the Fraser River Delta. Can. J. Plant. Sci.
Evans Ogden, L.J., K.A. Hobson, D.B. Lank and S. Bittman. 2005. Stable isotope analysis reveals that agricultural habitat provides an important dietary component for nonbreeding Dunlin. Avian Cons. Ecol. 1: (1):3(online)URL:http://www.ace-eco.org/vol1/iss1/art3/.
Evans Ogden, L.J., K.A. Hobson and D.B. Lank. 2004. Blood isotopic (d13C and d15N) turnover and diet-tissue fractionation factors in captive dunlin (Calidris alpina pacifica). Auk 121: 170-177.
Shepherd, P.C.F. and D.B. Lank. 2004. Marine and agricultural habitat preferences of dunlin wintering in British Columbia. J. Wildl. Manage. 68: 61-73.
Butler, R.W., P.C.F. Shepherd and M.J.F. Lemon. 2002. Site fidelity and local movements of migrating Western Sandpipers on the Fraser River estuary. Wilson Bull. 114: 485-490.
Evans Ogden, L. 2002. Non-breeding shorebirds in a coastal agricultural landscape: winter habitat use and dietary sources. PhD, Simon Fraser University, Burnaby, BC. 241 pp.
Shepherd, P. 2001. Status and conservation of Dunlin in Canada. Bird Trends 8:31-33.
Shepherd, P.C.F. 2001. Space use, habitat preferences, and time activity budgets of Dunlin (Calidris alpina pacifica) in the Fraser River Delta, BC. PhD, Simon Fraser University, Burnaby, BC. 204 pp.
Butler, R.W. 1999. Winter abundance and distribution of shorebirds and songbirds on farmlands on the Fraser River delta, British Columbia, 1989-1991. Can. Field Nat. 113: 390-395.
Shepherd, P. 1997. The winter ecology of Dunlin (Calidris alpina pacifica) in the Fraser River Delta. Report to: Environment Canada, Fraser River Action Plan. Technical Report, pp.
Sewell, M.A. 1996. Detection of the impact of predation by migratory shorebirds: an experimental test in the Fraser River estuary, British Columbia (Canada). Mar. Ecol. Prog. Ser. 144: 23-40.

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Other Shorebird Studies
Lozano, G.A. and D.B. Lank. 2004. Immunocompetence and testosterone dependent condition traits in male ruffs (Philomachus pugnax). Anim. Biol. 54: 315-329.
Schamel, D., D.M. Tracy and D.B. Lank. 2004. Male mate choice, male availability and egg production as limitations on polyandry in the Red-necked Phalarope. Anim. Behav. 67: 847-853.
Schamel, D., D.M. Tracy, D.B. Lank and D.F. Westneat. 2004. Copulation strategies and extra-pair fertilization rates in the sex-role reversed, socially polyandrous Red-necked Phalarope Phalaropus lobatus. Beh. Ecol. Sociobiol. 57: 110-118.
Zharikov, Y. and G.A. Skilleter. 2004. A relationship between prey density and territory size in non-breeding Eastern Curlews Numenius madagasceriensis. Ibis 146: 518-521.
Lozano, G.A. and D.B. Lank. 2003. Seasonal trade-offs in cell-mediated immunosenescence in ruffs (Philomachus pugnax). Proc. Roy. Soc. Lond. B. 270: 1203-1208.
Zharikov, Y. and G.A. Skilleter. 2003. Depletion of benthic invertebrates by bar-tailed godwits Limosa lapponica in a subtropical estuary. Mar. Ecol. Prog. Ser. 254: 151-162.
Zharikov, Y. and G.A. Skilleter. 2003. Nonbreeding eastern curlews Numenius madagascarienisis do not increase the rate of intake or digestive efficiency before long-distance migration because of an apparent digestive constraint. Physiol. Biochem. Zool. 76: 704-715.
Hazlitt, S. 2002. Territory quality and reproductive performance of Black Oystercatchers in British Columbia. Wilson Bull. 113: 404-409.
Hazlitt, S.L., R.C. Ydenberg and D.B. Lank. 2002. Territory structure, parental provisioning, and chick growth in the Black Oystercatcher (Haematopus bachmani). Ardea 90: 219-227.
Lank, D.B. 2002. Diverse processes maintain plumage polymorphisms in birds. J. Avian Biol. 33: 327-330.
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